Characterization of three plant or fungal feeding nematodes (Nematoda, Rhabditida, Tylenchomorpha) from coastal dunes in Spain including the SEM study of these species

Abstract Three known species, Aphelenchus avenae , Ditylenchus myceliophagus and Tylenchorhynchus aduncus (Rhabditida, Tylenchomorpha), are described from sand coastal dunes in Spain associated to the rhizosphere of xerophilic plants. A. is characterized by having stylet with 16–22 µm long, pharyngeal dorsal gland overlapping intestine, postvulval uterine sac about 2.5 times body diameter, female tail short conoid rounded tip bearing phasmid male bursa well developed. D. lateral field four incisures, 7–8 basal bulb irregularly pyriform, both sexes similar reaching three-quarters length. T. its six 15–18 μm spicules bifurcate distal ventral crest, gubernaculum hook-like manubrium. New morphological, including SEM study, morphometrical data included for three species. Also, phylogenetic analysis based on 18S 28S rDNA two species studied. Additionally, belonging superfamily Aphelenchoidea transferred other genera: assamensis genus Aphelenchoides dhanachandi Potensaphelenchus .